Hopelessly outmatched financially against Gryzzl in her attempt to acquire the huge Newport parcel (25 square miles of undeveloped land outside of town), Leslie hits on an idea: if she can have the Pawnee City Council declare that the land cannot be developed commercially, her dream of it becoming a new national park just might come true. Brilliant idea, but there's a huge problem: a preliminary tally shows the council is split two votes for each side. And that means the deciding vote is in the hands of an old adversary, Councilman Jeremy Jamm.
Leslie swallows her pride and heads to Jamm's office to ask for his support. But Jamm stops her halfway through her pitch; before any vote, he'll need to consult his girlfriend. To the horror of Leslie (and Parks fans everywhere), Jamm's new lady is none other than Tammy 2, Ron's evil ex-wife.
Leslie makes her pitch anyway, citing the open space and recreational use and the legacy a new park would create. Jamm reiterates that he'll vote however Tammy tells him to. When Tammy explains that she's only returned to town to destroy Ron Swanson, Leslie sees her opportunity. Her pitch is simple: by preventing Gryzzl and Ron from successfully developing the land, Tammy can crush him. It's a very tempting thought.
Meanwhile Tom is pouring out his soul to Andy at the bistro. He explains that he's lonely, ready to settle down and share his awesomeness with someone. Over a few drinks, Andy explains that a soulmate is out there waiting for Tom somewhere. They review Tom's Gryzzl profile and notice a new message from Lucy, a girl Tom used to date. Andy's convinced it's a sign. OK, so Lucy's in Chicago, but that's no impediment. Tom and Andy hop in a cab.
Meanwhile back at City Hall, April is incredibly excited: Ben has invited her to join him today when April's very favorite person on TV, Ms. Joan Callamezzo, comes by as part of the Pawnee Bicentennial Celebration. April gushes over the TV personality, asking her to sign her latest memoir, "Genius A'Flame: Joan Callamezzo: A Portrait in Words: Game of Joans."
In her prepared remarks, Joan explains that the reason she's so successful is because she's doing what's she's wanted to do since she was ten years old: be on TV. The secret to true happiness in life is doing what you love, she tells the crowd. The observation hits April like a brick. What is she doing with her own life?
Ben helps her navigate the career crisis. April tries a vocational aptitude test but it comes up with no matches. Finally, Ben asks April what she dreamed about doing when she was ten. A light bulb goes off and soon Ben and April are taking a tour of the local funeral home. As undertaker Dan Candle shows them around, Ben looks increasingly queasy while April grows more thrilled every minute. Could this be the start of her dream career?
Tammy pulls Leslie aside and informs her there's no chance she'll tell Jamm to vote for Leslie's proposal for non-development. Tammy says the real reason she's returned to Pawnee is to win Ron back.
Leslie bursts into Ron's office, declaring a truce. She pleads with Ron to help her break up Jamm and Tammy. Ron suspects the appeal is so that Jamm will vote for Leslie's proposal, but Leslie explains it's because she fears for Jamm himself. He's completely under Tammy's evil spell. Ron knows the sick, all-destructive power his ex has and agrees to join Leslie to help de-program Jamm.
They grab the councilman and secretly drive him up to Ron's remote cabin where they begin a painful regimen of aversion therapy. Exposing Jamm to Tammy's perfume, then slapping him silly, the tough love process is somewhere between a heroin addict's withdrawal and an exorcism. Again and again they ply him with pain, then practice various scenarios in which Tammy might put her clutches on him when they return to civilization. It's a heroic act: Ron and Leslie setting aside their differences to save the life of a man they both despise.
In Chicago, Andy and Tom stand outside the building where Lucy works. When she happens by, Tom's tongue-tied. Andy steps in, explaining that they're visiting town because he's moving there. Lucy agrees to meet up with them after she finishes up some work. Tom's obviously taken with Lucy, but afraid to make a move. With some encouragement from Andy, he finally agrees to ask Lucy if she'd like to come work for him back in Pawnee. To everyone's amazement, she says she'll seriously consider it. Of course, she'll have to ask her boyfriend first. Tom's tells her great. And in a few weeks, she actually shows up in Pawnee to start work. We'll see how this one shakes out.
Back at the funeral home, April's excited to see where the bodies are storied and prepared. But her giddiness evaporates when the undertaker explains that there are years of training required before anyone is allowed to process bodies for burial. April's dream of becoming a mortician dies in the cadaver preparation room.
She returns to her job and laments to Donna that she just can't quite find a career she's passionate about. Donna points out to April that she's not even 29 years old yet, which means that astrologically her Saturn return is still ahead of her. Every 29 years, Donna explains, the sun and Saturn realign on your birthday - and that's when life suddenly opens up with self-discovery. The answers are still ahead.
Leslie and Ron bring Jamm back to Pawnee's public library where they confront Tammy. Jamm steps forward and tells Tammy in a stern voice that they're breaking up. Tammy kicks into gear, plying Jeremy with baby talk and trying to expose Leslie's de-programming efforts as pure political manipulation. Jamm stands strong.
Frustrated, Tammy plays her final card: she strips naked, offering Jamm the heretofore forbidden fruit of her body. Encouraged by his former adversaries, Jamm is resolute, declaring to Tammy that the relationship is over. He walks out of the library with Leslie and Ron, leaving Tammy behind, naked, screaming and knocking over stands of books.
Back at council chambers, Jamm is his old, obnoxious self. He tells Ron and Leslie that he's abstaining from the zoning vote. Leslie realizes the vote will stall at two and two, making the whole procedure a waste of time. She and Ron exchange some reluctant compliments for each other over their bonding together to save Jamm. But soon enough, they're sworn enemies again.