It's Jane's birthday! Ben and Billy plan a huge surprise for her, but they let Jane think they've completely forgotten her big day, which, of course, really upsets Jane. Meanwhile, Ben intercepts a birthday card for Jane from their mom who left them years ago. Rita advises him to give it to her, but Ben doesn't want to. He wants to protect Jane from their mother.
At Donovan Decker, Jane fears her secret has been uncovered. (Little does she know that her co-workers just found out it's her birthday.) She manages to get her mind off of all of her troubles when Eli talks her into running errands with him around New York City. Jane ends up having a blast with Eli! When they return to Donovan Decker, she is stunned to find everyone (including Gray!) waiting for her with a birthday cake. Jane's excitement about her birthday being acknowledged is short-lived when she realizes Eli was just distracting her so her co-workers could set up the party. Meanwhile, Jeremy drinks too much at the party after answering to India's blackmail demands all day. In the end, he confesses that he was the mole to Jane!
Ben and Billy get worried when Jane never shows up to the surprise party they planned. They try texting her and calling, but Jane thinks her family and friends have forgotten her birthday, so she's celebrating with her co-workers. Eventually Ben goes to Donovan Decker and finds her. He admits to his sister that they knew it was her birthday all along but were just trying to surprise her. When they head back to the party, everyone is gone. But the surprises aren't over yet! This time both Jane AND Ben are surprised. Their long-lost mom is there!
Also, Jane meets Zoe. Nick has the opportunity to pitch in front of a big baseball scout. Amanda Clarke, blasts from Ben's past, returns to town - much to Rita's detest. And Zoe lies to Billy.
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