Nora, now in Monroe's custody, is given a dainty white dress in her prison cell and ordered to put it on. Monroe has prepared a nice meal for Nora, complete with Miles' favorite scotch. As she reluctantly sits at the table, he sweet-talks her in an attempt to extract Miles' current location. This is Monroe's last olive branch to Nora before things will get very bad for her. Instead of cracking, Nora grabs the scotch bottle and swings it at Monroe. She misses, but the message hits its target. Monroe orders his guards to take Nora away, and for the next 21 days, she's tortured incessantly. Where is Miles?!
On the 21st day of her torture, after multiple water boardings and injections of a hallucinogenic truth serum, Nora finally cracks. She tells Sanborn, who's been overseeing the sessions, where Miles is, also divulging Rachel's plan to reach The Tower in Colorado. When Monroe hears about The Tower, he brings Randall in for questioning, leveraging Sanborn's intel against him. Power is Monroe's only advantage in this war. Incensed that Randall kept him in the dark about The Tower, Monroe orders him killed. Claiming he's the only one who can get Monroe into The Tower, Randall wins his stay of execution, promising turning the lights back on is the least he can do. The Tower was a skunk works for the DOD; there are tools and technology there that even the President didn't know about. When Sanborn confirms Randall's story, Monroe decides it's time to get in the choppers and take a road trip to Colorado.
Rachel and Aaron finally reach Colorado in the pouring rain; unbeknownst to them, Monroe and his top advisors are racing to the same spot, determined to stop them. In Atlanta, Jason tells Charlie they should flee to Texas - he's really worried the city will be under siege any day now, and if Charlie gets caught, there's no telling what the Militia would do to her. They kiss in the middle of the street - Jason's falling for Charlie. Suddenly, Jason notices a man in the distance, Billings, and his demeanor changes. Something about that guy's presence makes Jason uneasy...
John Sanborn flees Monroe's compound with a drugged Nora, and brings her to Atlanta. When Miles hears of her return, he rushes to the hospital; Nora's going to be fine, but the drugs Monroe used on her are still in effect. Nevertheless, she tells Miles about her cracking and that Rachel is in danger. Miles interrogates Sanborn and comes to the same conclusion Monroe did earlier - Miles has to get to The Tower. Together with Neville, Hudson, Charlie and Jason, Miles loads up a Georgian helicopter and orders Sanborn to navigate. Meanwhile, in Colorado Springs, Monroe escorts Randall in chains to the entrance of The Tower, where the security touchpad denies Randall's handprint. Inside, a group of people watches the encounter on security monitors...
Jason holds Charlie's hand as the group flies towards Colorado; she's scared, as it's her first time flying. He flashes back to Atlanta and his conversation with Billings, who says he has a job for Jason. The helicopter has to land somewhere in the Plains Nation to refuel, but when they do, someone murders the helicopter's pilot, cutting his throat with a distinctive X, and messes with the vehicle's wiring. When Miles and Charlie go looking for the killers, they find Rebel commander Ramsey on the ground, bleeding out; he has the same distinctive wound on his neck. Miles tries to keep Ramsey alive long enough to figure out who's committing these acts, but Ramsey can't muster the words.
After reaching The Tower, Aaron and Rachel hide out in the woods, as Monroe's men are camped everywhere in front of the facility. Aaron doesn't know how they're going to access The Tower, but Rachel has a plan; that night, she'll kill Monroe, and in the ensuing chaos, Aaron will make a run for it, using Dr. Warren's notebook to access the door. Aaron hates this idea - there has to be a better way! If Rachel kills Monroe, his men will kill her in an instant. But Rachel knows Aaron won't get inside The Tower unless she does something drastic, and her goal all along was to get revenge for Danny and Ben. It's a sacrifice she's willing to make. It's up to Aaron now; he has to turn the power back on.
With no one around, Miles realizes the murderer has to be among their own group. Nora's nowhere to be found, so Miles orders everyone to place their weapons into a bag. As suspicions run hot, Miles throws the bag into the helicopter except for his own, then heads out to search for Nora. Finding her face down on the ground with a bloody arm and a knife, Miles fears the worst, but Nora's fine - though still hallucinating from Sanborn's drugs. Everyone argues Nora was the one who carried out the killings, though Miles insists it's not true. But why else would she be covered in blood with a knife next to her? Nora has to admit the group might be right...
Even though she admits to strangling a guard under Monroe's detainment while under the influence of Sanborn's hallucinogens, Miles insists on her innocence. Miles sifts through the bag of weapons to check for blood, until he notices Jason fumbling with something in his pocket - a bloody switchblade. Jason immediately says someone planted that on him. When he asks Charlie to back him up, she won't - who was the guy he was talking to back in Atlanta? After all, Jason has lied to her before. Jason admits that Billings, a Militia spy, asked him to kill Miles in exchange for Charlie's safety, but Jason said no. Miles doesn't believe him. When he pulls a gun on Jason, Charlie and Neville rush to his defense, giving Jason the opening to take off running.
Miles searches for Jason to no avail and is eventually stopped by Sanborn, who tells Miles the knife couldn't possibly be Jason's. It has the initials AMD stamped on it, which stands for Annapolis, Maryland, where the knife was made. Has anyone in their group been to Annapolis lately? Suddenly, a gunshot rings out and Sanborn goes down; Miles turns around to see Hudson holding the smoking gun! Hudson is the murderer!
Miles asks Hudson why he did it. Hudson tells Miles the Militia has his wife, and they're threatening to kill her unless he works for them; he was the one who gave Monroe Miles' location at Ashland, prompting the drone strike, and he knows killing Miles, Neville and the rest of their group - all high-value targets - would mean safety for his wife. With no other choice, Miles and Hudson fight to the death, a brutal, brawling bout that tests the limits of both men. But just as it seems as if Hudson has gained the upper hand, Jason comes out of nowhere and shoots him with Miles' discarded gun, which he then drops at Miles' feet. As Nora finishes fixing the helicopter, Charlie apologizes to Jason, who claims he gets it. He's lied to Charlie too many times - why should she trust him? Miles climbs into the driver's seat and the group resumes their journey to Colorado.
Meanwhile, at the Tower, Rachel kills a Militia guard and steals his uniform. That night, she walks through the camp in disguise, determined to find Monroe. Once she locates his tent, Rachel walks in and greets Monroe calmly. As his guards quickly train their weapons on her, Rachel calmly pulls the pin of a live grenade and simply smiles at Monroe. Her hand is still on the release valve as everyone in the room cautiously takes a step toward her.