With Commander Adama fighting for his life after being shot by the Cylon infiltrator Sharon, and President Roslin languishing in the Galactica's brig after losing the power struggle with Adama, Col. Tigh is thrust into the unfamiliar role of the sole leader in a time of crisis.
The situation worsens quickly. First, Tigh sends Lee to the brig for siding with Roslin against Commander Adama. Then an emergency jump to escape an incoming Cylon force goes wrong, leaving Galactica alone in space, separated from the rest of the fleet - and from the medical help needed to save Adama's life.
As Tigh struggles to come to terms with this crisis, he thinks back on his relationship with Adama, who saved him from oblivion and from his own fierce temper, and tries to tap into some of the wisdom that his old friend offered him.
It soon becomes clear that to reconnect with the fleet, Galactica must jump back into danger over Kobol and network its computers, a highly risky move that will make the ship vulnerable to attack from both the Cylon fighters and their crippling computer viruses.
Meanwhile, on the surface of Kobol, Chief Tyrol and the rest of the downed Raptor crew fight for their lives against an unseen enemy while Vice President Baltar seeks solace in his visions of Number Six, only to find those visions turn dark and haunting.
Light-years away, on Cylon-occupied Caprica, Kara and Helo must find some way to bring the Arrow of Apollo back to the fleet - but first they have to get it back from the avatar of Number Six who has found it in the Delphi Museum.