As the crew on Destiny are working around the clock to recover Scott, Chloe, Eli and Greer, several power systems on the ship cut out. Crew members clash over why this could be, but the rescue effort continues. TJ, continuing her psych evaluations of the crew, learns that James, insecure about her rating in Young's eyes, is also not yet over her breakup with Scott. TJ attempts to tell Young about her pregnancy, but her nerve fails her at the last minute.
In the underground tunnels of the alien planet, Eli has come up with a risky plan to get the team back to Destiny: Since they know Destiny's general trajectory, they can hopscotch from one planet to the next until they are in range of the ship - they just have to hope that they don't jump in the wrong direction. But unbeknownst to Eli, Rush has discovered an unnerving fact: Destiny is reaching the edge of a galaxy. This explains the power failures, but also means that the next time they drop out of FTL is their last and only chance to recover the recon team.
Chloe finally decodes a map underground, and just as the team discovers the exit, a tremor hits and Greer is caught behind a rockslide, apparently dead. With walls crumbling around them, the other three have no choice but to leave him to save themselves.
As Destiny drops out of FTL, Rush coordinates the same plan that Eli devised: Destiny crew members will check all gates in range in hopes of being close enough to hopscotch to the planet crew. Both the planet and ship crews begin the search, as Greer recovers consciousness and calls upon his childhood fears and traumas to motivate himself to dig out. He races to catch up with his team, but he is too late; they have taken the remote and left the planet. Alone, he is haunted by memories of abandonment, recalling his father's torment and his own heartbreaking decision to enlist in service.
Amid the chaos of the rescue mission, TJ finds a moment alone with Young to tell him she's carrying his child. He is visibly upset, but assures her that they'll make it work somehow.
Eli delivers good and bad news to Chloe and Scott: Although they've been jumping in the wrong direction, they are in range of the planet where Rush was left behind. Eli thinks he could find information there that will help them get back to Destiny, and although Chloe is against it, they investigate the alien shuttle on that planet. Eli can't make sense of it, but Chloe intuitively recognizes a star map. They head to the next planet, where Destiny's gate address comes into range. Meanwhile, the Destiny crew has found Greer. Both teams dial Destiny simultaneously and Rush, mere seconds ahead of Eli, steps through the gate and onto the ship. Eli is stumped as to why the dial didn't complete, tries again - and is again unsuccessful. As the ship crew take stock of their situation, Eli and the team realize that Destiny, so close to the edge of the galaxy, has jumped back into FTL, and out of range of the gate.