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Harris, who won a Tony recently for his role as the transgender German rocker, will not portray the role beyond August.
Harris, dressing up as Hedwig from his Broadway hit, performs 'Sugar Daddy' with co-star Lena Hall while Menzel wows the crowd with emotional performance of 'Always Starting Over' from 'If/Then'.
Bryan Cranston and Neil Patrick Harris predictably won the best performance by an actor in a play and musical respectively.
The actors took home one prize each but Harris had to share his with Jefferson Mays of 'A Gentleman's Guide to Love and Murder'.
Other performers for the awards show, which will air on CBS on June 8, include Sutton Foster, Gladys Knight, Patti LaBelle, Fantasia, Sting and Allan Cumming.
The CBS honcho clarifies Harris' statement that he was approached to replace David Letterman and says there are no talks with Joel McHale about the 12:30 slot.
The former 'How I Met Your Mother' star admits he was asked if he's interested in replacing David Letterman, but turned it down because he would get bored.
The actor, who is taking part in Broadway's 'Hedwig and the Angry Inch', only wears a black bowtie with a top hat covering his private parts.
This year's nomination list is dominated by 'A Gentleman's Guide to Love and Murder' with 10, followed by Harris' 'Hedwig and the Angry Inch' with 8.
A representative for the actor explains the whole sequence that actually happened during 'Hedwig and the Angry Inch'.
The 'How I Met Your Mother' star says the 'Late Show' gig would be 'an asinine amount of work,' but doesn't rule out the possibility of him taking over the show.
The 'How I Met Your Mother' co-stars sing 'Confrontation' from the musical as other stars including Alyson Hannigan, Cobie Smulders and Josh Radnor burst into lauhgter.
Joining the SLEEP with Neuro campaign, the actor sang a mid-tempo song while sitting on a bed scattered with rose petals.
The 'A Million Ways to Die in the West' star is feted in a drag-themed parade and roasted at Harvard's Farkas Hall.
Harvard University's Hasty Pudding Theatricals has chosen the 'How I Met Your Mother' star as Man of the Year, while Hellen Mirren was announced as Woman of the Year.